Insights from a profession grouse biologist on the components impacting jap ruffed grouse survival.
As an aspiring wildlife biologist, I used to be extraordinarily lucky to spend a semester of my undergraduate research in an internship program with Mr. William Okay. (Invoice) Igo, a biologist with the West Virginia Division of Pure Assets (WVDNR). Among the many many influences Invoice had on my life was his ardour for upland sport chook biology, administration, and searching.
Invoice and his Brittany gave me an important introduction to ruffed grouse searching. He sensed my curiosity and gave me a duplicate of The Upland Capturing Life by George Chook Evans. Invoice was fast to level out that Mr. Evans was considerably of an enigma for the WVDNR. He was a gifted author and fanatic for upland sport chook searching. Evans was a robust critic of late grouse seasons. He maintained that late season searching suppressed grouse populations.
Resulting from his notoriety, Mr. Evans had a whole lot of affect in his dwelling state of West Virginia. His writings had been repeated all through the area by outside writers and avid grouse hunters every time grouse populations fluctuated. It turned out that his ebook and principle of searching results on grouse formed my profession.
Preliminary Investigations of Virginia Ruffed Grouse
Following a pointy decline in grouse flushing charges in 1976, the Virginia Recreation Fee shortened the grouse season by two weeks. Mr. Joe Coggin, then-Supervisor of Recreation Analysis, instructed me George Chook attended the regulation hearings and spoke in favor of a shorter grouse season. However Joe was unsure late season searching was limiting Virginia grouse. He requested researchers at Virginia Tech to start an investigation.
I used to be lucky to be the primary Virginia Tech graduate pupil on this investigation. My advisor, Dr. Roy Kirkpatrick, directed me to start finding out the dietary ecology of grouse in Virginia. I particularly examined meals high quality and grouse situation. I discovered grouse had been capable of construct physique fat on diets of soppy mast in the summertime and fall, and acorns within the fall and winter. We hypothesized that winter survival and copy could possibly be impacted with out high-quality meals.

Years later, I grew to become Virginia’s Ruffed Grouse Undertaking Chief. The query of the impacts of late season grouse searching continued within the upland chook searching group. The basic query was whether or not late season searching was additive or compensatory mortality.


Is Late Season Ruffed Grouse Looking Additive Or Compensatory?
The premise for sport searching is that searching losses are compensated by fewer pure mortalities. Thus, post-hunting populations evaluate to unhunted populations. Wildlife managers additionally thought-about a 3rd speculation—that searching was compensatory as much as some threshold the place it then turns into additive. The plain concern was that searching would trigger a decline in breeding populations and end in long-term inhabitants declines.
Regrettably, wildlife research have concluded completely different outcomes on searching results on ruffed grouse populations. One clarification is that ruffed grouse ecology and hunter stress fluctuate extensively throughout its vary. In any case, it’s the most generally distributed resident sport chook in North America. Key components wildlife managers ideally need to know earlier than drawing any conclusions about searching impacts embody survival charges, searching mortality charges, and reproductive charges. Information of the impacts of climate, predation, illnesses, and habitats on all these parameters are necessary issues for managers. Thus, it’s comprehensible that completely different conclusions have been reported about searching and different components limiting grouse populations throughout the vary.
However, choices have to be made about setting seasons, with or with out knowledgeable data. Criticism and complaints from involved customers come when populations and harvests are lower than passable. Joe Coggin had felt the warmth. Now it was my flip as involved grouse hunters continued to boost the query about searching impacts.
Involved hunters needed to know if these hens harvested in late season would dwell and reproduce, thereby rising the inhabitants.
The Appalachian Cooperative Grouse Analysis Undertaking
Along with Jeff Sole, Upland Recreation Chook Undertaking Chief in Kentucky, we conceived a analysis mission to deal with these considerations. It took a number of years of planning and coordination, however we ultimately realized our wishful pondering of a cooperative multi-state analysis mission to analyze searching and different components limiting grouse populations.
The mission began small, however curiosity in grouse was excessive within the area. Ultimately, seven state wildlife businesses joined the mission. The mission was titled the Appalachian Cooperative Grouse Analysis Undertaking (ACGRP). Six years later, we had caught and radio collared 3,118 ruffed grouse at 12 research websites in eight states. A complete of 17 graduate college students at eight universities labored on the mission.
As a part of our major goal, we closed the searching seasons at three research websites over the mission’s last three years. We in contrast survival charges within the closed areas to 4 management websites the place searching continued into February.


We discovered no distinction in survival charges between hunted and un-hunted populations. In impact, we noticed that hens “saved” from searching had been however topic to excessive predation charges in March, April, and Might. The first spring predators had been hawks and owls. It is very important be aware that searching mortality charges in our research averaged 12 p.c.
Pat Devers, Ph.D. pupil at Virginia Tech engaged on the mission, wrote, “It is very important acknowledge that we can not assume harvest charges larger than these noticed on this research are compensatory; nor can we extrapolate our outcomes past the Appalachian area.”
We did discover searching elevated grouse winter dwelling ranges.


What Is Limiting Grouse Populations In The Appalachians?
Surprisingly, we discovered survival charges within the area had been akin to and even larger than research from the Lake States and New England. Many of the mortalities had been predation, with the best losses from avian predators, primarily within the fall and spring. We discovered Cooper’s hawks and owls had been the first predators.
Reproductive Charges
In distinction, we discovered reproductive charges within the area had been decrease. Most notably, chick survival charges and renesting charges had been decrease within the area’s most considerable forest sort, oak-hickory. We discovered important annual variation in copy in oak-hickory forests that correlated to acorn crops. We seen hens in these forests tended to forgo nesting within the spring following a poor acorn crop.
Physique Situation
Bob Lengthy, then-M.S. pupil at West Virginia College (WVU), discovered that diet and pre-breeding situation had a big influence on copy. Bob advised that hens have a threshold of roughly 11 p.c physique fats to achieve success nesting.
Climate And Predation
Past physique situation, climate and predation had been accountable for regulating chick survival charges. Utilizing miniature transmitters on grouse chicks, Brian Smith, then-Ph.D. candidate at WVU, discovered predation and climate had been equally accountable for chick losses.
Extra Findings From The ACGRP
In distinction to oak-hickory habitats, grouse in hardwood forests (cherry-maple) within the Appalachians have extra dependable meals assets and extra predictable copy. John Tirpak, then-M.S. pupil at California College of Pennsylvania, discovered nests in dense understories had been extra doubtless to achieve success than these in sapling stands and people close to openings.
Darroch Whitaker, then-Ph.D. pupil at Virginia Tech, studied the house ranges and house use of grouse throughout 10 research websites. He examined 67,814 areas of radio-marked grouse to find out dwelling ranges of greater than 1,000 birds. Darroch discovered that females had bigger dwelling ranges than males, juveniles had bigger dwelling ranges than adults, and hens with broods used moist bottomlands and riparian areas. He concluded managers ought to preserve and improve these essential areas to extend grouse copy and develop populations.


Japanese Ruffed Grouse Administration Immediately
The findings from the ACGRP underscored the complexity of ruffed grouse inhabitants dynamics within the Appalachian area. Whereas searching mortality was discovered to be largely compensatory beneath the noticed situations, the long-term viability of grouse populations is tied to broader ecological components, significantly habitat high quality and meals availability. Immediately, jap ruffed grouse administration efforts are more and more targeted on addressing these elementary challenges.
The “Japanese Ruffed Grouse Conservation Plan (2025-2034)” builds on these insights. The Plan outlines a regional technique that prioritizes habitat restoration, standardized monitoring, and collaborative conservation efforts throughout a number of states and provinces.
Finally, the way forward for ruffed grouse within the jap U.S. relies on a landscape-scale dedication to habitat restoration and adaptive administration. Science-driven conservation methods make sure that administration choices are primarily based on a complete understanding of the species’ wants. By implementing the methods outlined within the Plan, wildlife managers goal to stabilize and, the place potential, rebuild grouse populations for future generations.
“And for as long as the gunner treasures the birds of a paradise and treats them as the delicate issues they’re,” wrote George Chook Evans, “every covet will belong to him and his canine. Or so he could dream.”
Learn Extra
Trial by Fireplace: The Loss and Potential Restoration of Ruffed Grouse Habitat in Virginia
Ruffed Grouse Drumming – The Mysterious Noise of Northern Forests
Views – A West Virginia Looking Movie
Understanding the Risk Response of Ruffed Grouse in Looking